When in late 1938 the DB 601 engine, which used fuel injection system and not the usual carburetor, became available the Bf.110C was put into production, so in early 1939 the first pre-production version, designated as Bf.110C-0 was supplied to some units of the Luftwaffe. Of that C-0 variant only ten machines were built but were followed by the C-1, with almost 160 machines manufactured by 1st September 1939.
The DB 601 engines changed the external appearance of the aircraft, especially the engine nacelles, which were now faired due to the main radiator.
Wing tips were changed too, as they were no longer rounded, but square. This change was kept on every later Bf.110 until the end of the war, as it reduced the wingspan and wing surface, making it more manoeuvrable.
When the war broke out in September 1939 less than one hundred Bf.100C-1s were operational. Of those, eighty were equipping three units: I.(Zerstörer)/LG 1 , which was an operational development unit, I./ZG 1 and I./ZG 76. There seven additional Zerstörer squadrons, but they were still equipped with Messerschmitt Bf.109E fighters due to the lack of Bf.110s.
As the Bf.110 appeared too late to take part in the Spanish Civil War, it was employed first during the Polish Campaign. During this time only I.(Z)/LG 1, based at Greifswald, was fully equipped with the Bf.110C, since the two other groups were still operating the Jumo-powered Bf.110B. The aforementioned three units were immediately put into action by escorting bombers or strafing ground targets, tasks which they carried out with great efficiency with a total of 12 machines lost, reinforcing Luftwaffe's reputation of being invincible.
The success of the Polish Campaign and the good operational numbers shown by the Bf.110, prompted further development of the Bf.110C.
This way the Bf.110C-2 was identical to C-1 but had better radio equipment, the Bf.110C-3 was equipped with MG FF/M cannons, which became standard on the following variants, the Bf.110C-4 incorporated every mentioned previous improvement and featured also extra armour, making it heavier. The C-4 was also equipped with bomb racks allowing it to carry two 250 kg (550 lb) bombs and new engines were fitted in order to make it more powerful; the DB 601Ns rated at 1,270 hp.
The Bf.110C-5 was a reconnaissance variant which got the ventral cannon replaced by a photographic camera, the C-6 was a semi-experimental version of which only twelve machines were made and was equipped with a ventral-mounted 30 mm Mk 101 cannon replacing the original 20 mm one. The C-6 was put into service in the spring of 1940.
The Bf.110C-7 was a Jabo (Jagdbomber - Fighter-Bomber) variant with reinforced undercarriage to carry two 500 kg (1,100 lb) bombs placed side by side under the fuselage. This variant was manufactured in limited numbers as the C-4 was the main production version.
After the end of the Polish Campaign, the during the period of the Phoney War the Zerstörer activity was limited, although they were redeployed to the Western frontier of Germany.
The first aerial claim of the Bf.110 took place on 23rd November 1939 when various Bf.110s of LG 1 engaged and shot down a French Morane-Saulnier Ms.406 over Verdun. Three week later, on 18th December, one the first major engagements of the war took place, when the British Royal Air Force (RAF) sent 22 Vickers Wellington bombers to attack the German naval base at Wilhelmshaven. The British bombers could not find their targets and turned back... to be intercepted by a formation of sixteen Bf.110Cs belonging to I./ZG 76 and thirty-four Bf.109s. In spite of the help given by the Bf.109s, it was the Bf.110 which showed its great capacity as a bomber destroyer and, by the end of the fight, official Luftwaffe reports counted 38 British bombers shot down, a number clearly exaggerated as actual losses were 11 Wellingtons shot down and six damaged, with some sources claiming a 12th Wellington as destroyed. This disastrous raid convinced the RAF's Bomber Command to abandon day bombing in favour of night raids.
Sources:
1st Histoire & Collections - Avions et Pilotes 11 - Messerschmitt Bf.110 & Me.210-410 1939 -1945
2nd https://en.wikipedia.org/wiki/Messerschmitt_Bf_110_operational_history
3rd https://www.armedconflicts.com/prehled-verzi-t15722
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